Among animals, symbols are transmitted by tradition from generation to generation, and it is here, if one wishes, that one may draw the border line between the "animal" and man. In animals, individually acquired experience is sometimes transmitted by teaching and learning, from elder to younger individuals, though such true tradition is only seen in those forms whose high capacity for learning is combined with a higher development of their social life. True tradition has been demonstrated in jackdaws, greyleg geese, and rats. But knowledge thus transmitted is limited to very simple things, such as pathfinding, recognition of certain foods and of enemies of the species, and—in rats—knowledge of the danger of poisons. However, no means of communication, no learned rituals are ever handed by tradition in animals. In other words, animals have no culture.